Professor Z

Professor Zündapp, better known as Professor Z and also known as the Professor, is the secondary antagonist of Cars 2. He is the sidekick of Miles Axlerod, and the leader of the Lemon Cars. He is the main villain.

History
Professor Z is an internationally wanted weapons designer in a small, sophisticated German package. He is a brilliant, but mad monocle-wearing scientist and the leader of criminals called the Lemon Cars, who are plotting to sabotage the World Grand Prix.

Though his true motive is unclear, Professor Z is willing to do whatever it takes to eliminate all obstacles and keep the "project" on schedule. He is also responsible for inventing elector-magnetic pulse weapons designed as World Grand Prix cameras, intending to ignite Allinol, the 'alternative energy' fuel used by all race cars of the Grand Prix.

During the climax of the film, Professor Z orders his thugs to kill Lightning McQueen for the final step of their plans since Lightning decides to remain using Allinol for the final race in England, despite the fact that Allinol is being called into question.

However, in the final race in England, the Allinol that Lightning's using was actually replaced by Fillmore's organic fuel mixture that prevents the radiation from killing him. As a backup plan, Professor Z implants a bomb on Mater's air filter to make sure that Lightning dies anyway, but he is unable to detonate the bomb as it went out of range since Mater tries his best to lure everyone away from its threat while Lightning follows him unaware of the situation. Professor Z then send the lemon cars to surround Lightning and Mater in an intersection to kill them, but they are foiled by the combined efforts of British agents Finn McMissile and Holley Shiftwell and the Radiator Springs residents. Professor Z ends up being tied up by Finn while trying to escape and later gets shocked by Holley before getting arrested. Also, Mater points out that Allinol was actually gasoline engineered to ignite if hit with the EMP beam in order to discredit real alternative energy. As he finally reveals the story of the Prix conspiracy to the public by confronting and forcing Miles Axlerod to deactivate the bomb with a voice command to prevent being killed by it, Profesor Z, Miles, and the Lemon Cars are presumed to be thrown into prison for good.